Gpr17 (G protein-coupled receptor 17) - Rat Genome Database

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Gene: Gpr17 (G protein-coupled receptor 17) Rattus norvegicus
Analyze
Symbol: Gpr17
Name: G protein-coupled receptor 17
RGD ID: 1589785
Description: Enables receptor serine/threonine kinase binding activity. Involved in oligodendrocyte differentiation. Predicted to be active in plasma membrane. Orthologous to human GPR17 (G protein-coupled receptor 17); INTERACTS WITH 2,3,7,8-tetrachlorodibenzodioxine; bisphenol A; diethylstilbestrol.
Type: protein-coding
RefSeq Status: VALIDATED
Previously known as: G-protein coupled receptor 17; UDP/CysLT receptor; uracil nucleotide/cysteinyl leukotriene receptor
RGD Orthologs
Human
Mouse
Chinchilla
Bonobo
Dog
Squirrel
Pig
Green Monkey
Naked Mole-Rat
Alliance Orthologs
More Info more info ...
Latest Assembly: mRatBN7.2 - mRatBN7.2 Assembly
Position:
Rat AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
GRCr81823,850,460 - 23,857,381 (-)NCBIGRCr8
mRatBN7.21823,576,232 - 23,583,153 (-)NCBImRatBN7.2mRatBN7.2
mRatBN7.2 Ensembl1823,577,242 - 23,582,966 (-)EnsemblmRatBN7.2 Ensembl
UTH_Rnor_SHR_Utx1823,708,403 - 23,709,422 (-)NCBIRnor_SHRUTH_Rnor_SHR_Utx
UTH_Rnor_SHRSP_BbbUtx_1.01824,457,393 - 24,458,412 (-)NCBIRnor_SHRSPUTH_Rnor_SHRSP_BbbUtx_1.0
UTH_Rnor_WKY_Bbb_1.01823,803,373 - 23,804,392 (-)NCBIRnor_WKYUTH_Rnor_WKY_Bbb_1.0
Rnor_6.01824,733,306 - 24,739,194 (-)NCBIRnor6.0Rnor_6.0rn6Rnor6.0
Rnor_6.0 Ensembl1824,734,330 - 24,735,349 (-)EnsemblRnor6.0rn6Rnor6.0
Rnor_5.01824,446,883 - 24,453,499 (-)NCBIRnor5.0Rnor_5.0rn5Rnor5.0
RGSC_v3.41824,374,253 - 24,375,272 (-)NCBIRGSC3.4RGSC_v3.4rn4RGSC3.4
Celera1823,332,880 - 23,333,899 (-)NCBICelera
Cytogenetic Map18p12NCBI
JBrowse: View Region in Genome Browser (JBrowse)
Model


Gene Ontology Annotations     Click to see Annotation Detail View

Cellular Component

Molecular Function

References

References - curated
# Reference Title Reference Citation
1. Does GRK-ß arrestin machinery work as a "switch on" for GPR17-mediated activation of intracellular signaling pathways? Daniele S, etal., Cell Signal. 2014 Jun;26(6):1310-25. doi: 10.1016/j.cellsig.2014.02.016. Epub 2014 Mar 5.
2. The ubiquitin ligase Mdm2 controls oligodendrocyte maturation by intertwining mTOR with G protein-coupled receptor kinase 2 in the regulation of GPR17 receptor desensitization. Fumagalli M, etal., Glia. 2015 Dec;63(12):2327-39. doi: 10.1002/glia.22896. Epub 2015 Jul 31.
3. Phylogenetic-based propagation of functional annotations within the Gene Ontology consortium. Gaudet P, etal., Brief Bioinform. 2011 Sep;12(5):449-62. doi: 10.1093/bib/bbr042. Epub 2011 Aug 27.
4. GOA pipeline RGD automated data pipeline
5. Data Import for Chemical-Gene Interactions RGD automated import pipeline for gene-chemical interactions
Additional References at PubMed
PMID:16990797   PMID:20056144   PMID:20574000   PMID:21209081   PMID:22155652   PMID:23288840   PMID:24150254   PMID:26506265   PMID:26620557   PMID:27544384   PMID:28254957   PMID:29187582  
PMID:30226562   PMID:32387084   PMID:34569901   PMID:34769111  


Genomics

Comparative Map Data
Gpr17
(Rattus norvegicus - Norway rat)
Rat AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
GRCr81823,850,460 - 23,857,381 (-)NCBIGRCr8
mRatBN7.21823,576,232 - 23,583,153 (-)NCBImRatBN7.2mRatBN7.2
mRatBN7.2 Ensembl1823,577,242 - 23,582,966 (-)EnsemblmRatBN7.2 Ensembl
UTH_Rnor_SHR_Utx1823,708,403 - 23,709,422 (-)NCBIRnor_SHRUTH_Rnor_SHR_Utx
UTH_Rnor_SHRSP_BbbUtx_1.01824,457,393 - 24,458,412 (-)NCBIRnor_SHRSPUTH_Rnor_SHRSP_BbbUtx_1.0
UTH_Rnor_WKY_Bbb_1.01823,803,373 - 23,804,392 (-)NCBIRnor_WKYUTH_Rnor_WKY_Bbb_1.0
Rnor_6.01824,733,306 - 24,739,194 (-)NCBIRnor6.0Rnor_6.0rn6Rnor6.0
Rnor_6.0 Ensembl1824,734,330 - 24,735,349 (-)EnsemblRnor6.0rn6Rnor6.0
Rnor_5.01824,446,883 - 24,453,499 (-)NCBIRnor5.0Rnor_5.0rn5Rnor5.0
RGSC_v3.41824,374,253 - 24,375,272 (-)NCBIRGSC3.4RGSC_v3.4rn4RGSC3.4
Celera1823,332,880 - 23,333,899 (-)NCBICelera
Cytogenetic Map18p12NCBI
GPR17
(Homo sapiens - human)
Human AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
GRCh382127,646,153 - 127,652,639 (+)NCBIGRCh38GRCh38hg38GRCh38
GRCh38.p14 Ensembl2127,645,864 - 127,652,639 (+)EnsemblGRCh38hg38GRCh38
GRCh372128,403,728 - 128,410,213 (+)NCBIGRCh37GRCh37hg19GRCh37
Build 362128,120,217 - 128,126,683 (+)NCBINCBI36Build 36hg18NCBI36
Build 342128,119,976 - 128,126,442NCBI
Celera2121,714,385 - 121,721,160 (+)NCBICelera
Cytogenetic Map2q14.3NCBI
HuRef2120,709,795 - 120,716,458 (+)NCBIHuRef
CHM1_12128,407,871 - 128,414,621 (+)NCBICHM1_1
T2T-CHM13v2.02128,081,407 - 128,087,893 (+)NCBIT2T-CHM13v2.0
Gpr17
(Mus musculus - house mouse)
Mouse AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
GRCm391832,076,052 - 32,082,925 (-)NCBIGRCm39GRCm39mm39
GRCm39 Ensembl1832,076,050 - 32,082,689 (-)EnsemblGRCm39 Ensembl
GRCm381831,942,999 - 31,949,890 (-)NCBIGRCm38GRCm38mm10GRCm38
GRCm38.p6 Ensembl1831,942,997 - 31,949,636 (-)EnsemblGRCm38mm10GRCm38
MGSCv371832,102,653 - 32,109,290 (-)NCBIGRCm37MGSCv37mm9NCBIm37
MGSCv361832,086,024 - 32,092,646 (-)NCBIMGSCv36mm8
Celera1832,433,365 - 32,440,002 (-)NCBICelera
Cytogenetic Map18B1NCBI
cM Map1817.89NCBI
Gpr17
(Chinchilla lanigera - long-tailed chinchilla)
Chinchilla AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
ChiLan1.0 EnsemblNW_0049554592,890,266 - 2,903,053 (-)EnsemblChiLan1.0
ChiLan1.0NW_0049554592,890,266 - 2,903,053 (-)NCBIChiLan1.0ChiLan1.0
GPR17
(Pan paniscus - bonobo/pygmy chimpanzee)
Bonobo AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
NHGRI_mPanPan1-v21347,991,565 - 47,999,056 (-)NCBINHGRI_mPanPan1-v2
NHGRI_mPanPan12B48,006,543 - 48,013,269 (-)NCBINHGRI_mPanPan1
Mhudiblu_PPA_v02B32,623,444 - 32,630,256 (-)NCBIMhudiblu_PPA_v0Mhudiblu_PPA_v0panPan3
PanPan1.12B128,312,116 - 128,319,177 (+)NCBIpanpan1.1PanPan1.1panPan2
PanPan1.1 Ensembl2B128,316,464 - 128,321,519 (+)Ensemblpanpan1.1panPan2
GPR17
(Canis lupus familiaris - dog)
Dog AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
CanFam3.11922,958,778 - 22,968,489 (-)NCBICanFam3.1CanFam3.1canFam3CanFam3.1
CanFam3.1 Ensembl1922,963,363 - 22,964,382 (-)EnsemblCanFam3.1canFam3CanFam3.1
Dog10K_Boxer_Tasha1923,205,988 - 23,215,825 (-)NCBIDog10K_Boxer_Tasha
ROS_Cfam_1.01924,284,819 - 24,294,651 (-)NCBIROS_Cfam_1.0
ROS_Cfam_1.0 Ensembl1924,284,851 - 24,294,483 (-)EnsemblROS_Cfam_1.0 Ensembl
UMICH_Zoey_3.11922,988,450 - 22,998,293 (-)NCBIUMICH_Zoey_3.1
UNSW_CanFamBas_1.01923,183,202 - 23,193,039 (-)NCBIUNSW_CanFamBas_1.0
UU_Cfam_GSD_1.01924,315,107 - 24,324,942 (-)NCBIUU_Cfam_GSD_1.0
Gpr17
(Ictidomys tridecemlineatus - thirteen-lined ground squirrel)
Squirrel AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
HiC_Itri_2NW_02440530397,960,251 - 97,968,807 (+)NCBIHiC_Itri_2
SpeTri2.0 EnsemblNW_00493646943,653,481 - 43,654,497 (-)EnsemblSpeTri2.0SpeTri2.0 Ensembl
SpeTri2.0NW_00493646943,652,244 - 43,656,661 (-)NCBISpeTri2.0SpeTri2.0SpeTri2.0
GPR17
(Sus scrofa - pig)
Pig AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
Sscrofa11.1 Ensembl1559,692,996 - 59,701,265 (-)EnsemblSscrofa11.1susScr11Sscrofa11.1
Sscrofa11.11559,692,992 - 59,702,513 (-)NCBISscrofa11.1Sscrofa11.1susScr11Sscrofa11.1
Sscrofa10.21566,443,203 - 66,448,850 (+)NCBISscrofa10.2Sscrofa10.2susScr3
GPR17
(Chlorocebus sabaeus - green monkey)
Green Monkey AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
ChlSab1.1102,564,156 - 2,571,442 (-)NCBIChlSab1.1ChlSab1.1chlSab2
Vero_WHO_p1.0NW_02366606120,189,037 - 20,195,643 (+)NCBIVero_WHO_p1.0Vero_WHO_p1.0
Gpr17
(Heterocephalus glaber - naked mole-rat)
Naked Mole-Rat AssemblyChrPosition (strand)SourceGenome Browsers
JBrowseNCBIUCSCEnsembl
HetGla_female_1.0 EnsemblNW_00462473214,115,325 - 14,128,169 (+)EnsemblHetGla_female_1.0HetGla_female_1.0 EnsemblhetGla2
HetGla 1.0NW_00462473214,115,267 - 14,129,714 (+)NCBIHetGla_female_1.0HetGla 1.0hetGla2

Variants

.
Variants in Gpr17
15 total Variants
miRNA Target Status

Predicted Target Of
Summary Value
Count of predictions:57
Count of miRNA genes:51
Interacting mature miRNAs:56
Transcripts:ENSRNOT00000036537
Prediction methods:Miranda, Rnahybrid, Targetscan
Result types:miRGate_prediction

The detailed report is available here: Full Report CSV TAB Printer

miRNA Target Status data imported from miRGate (http://mirgate.bioinfo.cnio.es/).
For more information about miRGate, see PMID:25858286 or access the full paper here.


QTLs in Region (mRatBN7.2)
The following QTLs overlap with this region.    Full Report CSV TAB Printer Gviewer
RGD IDSymbolNameLODP ValueTraitSub TraitChrStartStopSpecies
2301410Bp317Blood pressure QTL 3170.004arterial blood pressure trait (VT:2000000)mean arterial blood pressure (CMO:0000009)181194427326548295Rat
1358358Sradr6Stress Responsive Adrenal Weight QTL 62.49adrenal gland mass (VT:0010420)both adrenal glands wet weight (CMO:0000164)181194429959330563Rat
1331735Rf44Renal function QTL 442.981urine total protein amount (VT:0000032)urine total protein excretion rate (CMO:0000756)181823456431359530Rat
61382Bp46Blood pressure QTL 4618.8arterial blood pressure trait (VT:2000000)systolic blood pressure (CMO:0000004)181194179131393320Rat
1331741Bp232Blood pressure QTL 2323.59112arterial blood pressure trait (VT:2000000)diastolic blood pressure (CMO:0000005)182137289383213037Rat
8552968Pigfal19Plasma insulin-like growth factor 1 level QTL 1911.4blood insulin-like growth factor amount (VT:0010479)plasma insulin-like growth factor 1 level (CMO:0001299)18125758124Rat
61388Bp2Blood pressure QTL 23.23arterial blood pressure trait (VT:2000000)diastolic blood pressure (CMO:0000005)18135374722Rat
1331781Scl28Serum cholesterol level QTL 283.995blood HDL cholesterol amount (VT:0000184)serum high density lipoprotein cholesterol level (CMO:0000361)181194429924797117Rat
6903359Bp355Blood pressure QTL 3553.6arterial blood pressure trait (VT:2000000)systolic blood pressure (CMO:0000004)181194454459796478Rat
2313082Bss85Bone structure and strength QTL 850.80.0001long bone metaphysis morphology trait (VT:0000133)tibia midshaft total cross-sectional area (CMO:0001715)181495133759951337Rat
1641923Colcr8Colorectal carcinoma resistance QTL 83.10.0014intestine integrity trait (VT:0010554)poorly differentiated malignant colorectal tumor number (CMO:0002076)182206624252293055Rat
2312568Glom21Glomerulus QTL 2120.005kidney glomerulus morphology trait (VT:0005325)index of glomerular damage (CMO:0001135)18518585840503530Rat
1331766Bp236Blood pressure QTL 2363.022arterial blood pressure trait (VT:2000000)mean arterial blood pressure (CMO:0000009)181194429931359530Rat
1358193Emca2Estrogen-induced mammary cancer QTL 21.6mammary gland integrity trait (VT:0010552)post-insult time to mammary tumor formation (CMO:0000345)181800759963571040Rat
631264Scl22Serum cholesterol level QTL 226.2blood cholesterol amount (VT:0000180)serum total cholesterol level (CMO:0000363)181194179131393320Rat
1331775Bp235Blood pressure QTL 2353.201arterial blood pressure trait (VT:2000000)mean arterial blood pressure (CMO:0000009)181823456431359530Rat
12904680Bw189Body weight QTL 1890.019body mass (VT:0001259)body weight (CMO:0000012)181194427326548295Rat
1600373Mamtr6Mammary tumor resistance QTL 6mammary gland integrity trait (VT:0010552)mammary tumor number (CMO:0000343)181927890183218561Rat
2293708Bss46Bone structure and strength QTL 468.80.0001lumbar vertebra morphology trait (VT:0010494)lumbar vertebra cortical cross-sectional area (CMO:0001690)181179151863636873Rat
12904693Am20Aortic mass QTL 200.001aorta mass (VT:0002845)aorta weight to aorta length to body weight ratio (CMO:0002722)181194427326548295Rat
9589153Insul31Insulin level QTL 317.150.05blood insulin amount (VT:0001560)plasma insulin level (CMO:0000342)18142547119Rat
1578667Bss21Bone structure and strength QTL 213.5femur morphology trait (VT:0000559)femoral neck cortical cross-sectional area (CMO:0001702)181179151883218561Rat
12904695Kidm73Kidney mass QTL 730.001kidney mass (VT:0002707)both kidneys wet weight to body weight ratio (CMO:0000340)181194427326548295Rat
2312598Bp340Blood pressure QTL 3400.05arterial blood pressure trait (VT:2000000)systolic blood pressure (CMO:0000004)18130558703Rat
12904689Cm128Cardiac mass QTL 1280.001heart mass (VT:0007028)heart wet weight to body weight ratio (CMO:0002408)181194427326548295Rat
12904690Cm129Cardiac mass QTL 1290.001heart left ventricle mass (VT:0007031)heart left ventricle weight to body weight ratio (CMO:0000530)181194427326548295Rat
2300180Bmd67Bone mineral density QTL 674.80.0001femur mineral mass (VT:0010011)bone mineral density (CMO:0001226)18134291613Rat
12904691Cm130Cardiac mass QTL 1300.001heart right ventricle mass (VT:0007033)heart right ventricle weight to body weight ratio (CMO:0000914)181194427326548295Rat
9590248Scort10Serum corticosterone level QTL 1019.710.001blood corticosterone amount (VT:0005345)plasma corticosterone level (CMO:0001173)18125758124Rat
2299160Iddm35Insulin dependent diabetes mellitus QTL 352.79blood glucose amount (VT:0000188)age at onset/diagnosis of type 1 diabetes mellitus (CMO:0001140)18420794160377792Rat
1578661Bss20Bone structure and strength QTL 203.7femur morphology trait (VT:0000559)femoral neck cross-sectional area (CMO:0001697)181179151883218561Rat
1331753Bp231Blood pressure QTL 2313.643arterial blood pressure trait (VT:2000000)systolic blood pressure (CMO:0000004)181194429952293055Rat
2293661Bss50Bone structure and strength QTL 504.640.0003lumbar vertebra size trait (VT:0010518)lumbar vertebra trabecular cross-sectional area (CMO:0001692)18134291613Rat
61375Bp41Blood pressure QTL 412.4blood pressure trait (VT:0000183)systolic blood pressure (CMO:0000004)181194429941122201Rat


Expression


RNA-SEQ Expression
High: > 1000 TPM value   Medium: Between 11 and 1000 TPM
Low: Between 0.5 and 10 TPM   Below Cutoff: < 0.5 TPM

alimentary part of gastrointestinal system circulatory system endocrine system exocrine system hemolymphoid system hepatobiliary system integumental system musculoskeletal system nervous system renal system reproductive system respiratory system appendage
High
Medium 3 130
Low 18 70 60 27 39 27 23 21 110 24 69 35 21
Below cutoff 3 26 82 38 27 38 13 21 68 58 3

Sequence


Ensembl Acc Id: ENSRNOT00000036537   ⟹   ENSRNOP00000032046
Type: CODING
Position:
Rat AssemblyChrPosition (strand)Source
Rnor_6.0 Ensembl1824,734,330 - 24,735,349 (-)Ensembl
Ensembl Acc Id: ENSRNOT00000104846   ⟹   ENSRNOP00000091710
Type: CODING
Position:
Rat AssemblyChrPosition (strand)Source
mRatBN7.2 Ensembl1823,577,242 - 23,582,966 (-)Ensembl
RefSeq Acc Id: NM_001071777   ⟹   NP_001065245
RefSeq Status: VALIDATED
Type: CODING
Position:
Rat AssemblyChrPosition (strand)Source
GRCr81823,850,460 - 23,857,381 (-)NCBI
mRatBN7.21823,576,232 - 23,583,153 (-)NCBI
Rnor_6.01824,734,330 - 24,735,349 (-)NCBI
Rnor_5.01824,446,883 - 24,453,499 (-)NCBI
RGSC_v3.41824,374,253 - 24,375,272 (-)RGD
Celera1823,332,880 - 23,333,899 (-)RGD
Sequence:
Protein Sequences
Protein RefSeqs NP_001065245 (Get FASTA)   NCBI Sequence Viewer  
GenBank Protein ABG75921 (Get FASTA)   NCBI Sequence Viewer  
  EDL76180 (Get FASTA)   NCBI Sequence Viewer  
Ensembl Protein ENSRNOP00000091710.1
  ENSRNOP00060039918
  ENSRNOP00065015342
GenBank Protein Q09QM4 (Get FASTA)   NCBI Sequence Viewer  
RefSeq Acc Id: NP_001065245   ⟸   NM_001071777
- UniProtKB: Q09QM4 (UniProtKB/Swiss-Prot),   A6J2P8 (UniProtKB/TrEMBL)
- Sequence:
Ensembl Acc Id: ENSRNOP00000032046   ⟸   ENSRNOT00000036537
Ensembl Acc Id: ENSRNOP00000091710   ⟸   ENSRNOT00000104846
Protein Domains
G-protein coupled receptors family 1 profile

Protein Structures
Name Modeler Protein Id AA Range Protein Structure
AF-Q09QM4-F1-model_v2 AlphaFold Q09QM4 1-339 view protein structure

Transcriptome

eQTL   View at Phenogen
WGCNA   View at Phenogen
Tissue/Strain Expression   View at Phenogen


Additional Information

Database Acc Id Source(s)
AGR Gene RGD:1589785 AgrOrtholog
BioCyc Gene G2FUF-8047 BioCyc
Ensembl Genes ENSRNOG00000025745 Ensembl
  ENSRNOG00000065480 Ensembl, UniProtKB/Swiss-Prot
  ENSRNOG00060027654 UniProtKB/Swiss-Prot
  ENSRNOG00065012288 UniProtKB/Swiss-Prot
Ensembl Transcript ENSRNOT00000104846.1 UniProtKB/Swiss-Prot
  ENSRNOT00060047974 UniProtKB/Swiss-Prot
  ENSRNOT00065019996 UniProtKB/Swiss-Prot
Gene3D-CATH Rhodopsin 7-helix transmembrane proteins UniProtKB/Swiss-Prot, UniProtKB/TrEMBL
InterPro GPCR_Rhodpsn UniProtKB/Swiss-Prot, UniProtKB/TrEMBL
  GPCR_Rhodpsn_7TM UniProtKB/Swiss-Prot, UniProtKB/TrEMBL
KEGG Report rno:767613 UniProtKB/Swiss-Prot, UniProtKB/TrEMBL
NCBI Gene 767613 ENTREZGENE
PANTHER G-PROTEIN COUPLED RECEPTOR UniProtKB/Swiss-Prot, UniProtKB/TrEMBL
  URACIL NUCLEOTIDE/CYSTEINYL LEUKOTRIENE RECEPTOR UniProtKB/Swiss-Prot, UniProtKB/TrEMBL
Pfam 7tm_1 UniProtKB/Swiss-Prot, UniProtKB/TrEMBL
PhenoGen Gpr17 PhenoGen
PRINTS GPCRRHODOPSN UniProtKB/Swiss-Prot, UniProtKB/TrEMBL
  P2YPURNOCPTR UniProtKB/Swiss-Prot, UniProtKB/TrEMBL
PROSITE G_PROTEIN_RECEP_F1_1 UniProtKB/Swiss-Prot, UniProtKB/TrEMBL
  G_PROTEIN_RECEP_F1_2 UniProtKB/Swiss-Prot, UniProtKB/TrEMBL
RatGTEx ENSRNOG00000025745 RatGTEx
  ENSRNOG00000065480 RatGTEx
  ENSRNOG00060027654 RatGTEx
  ENSRNOG00065012288 RatGTEx
Superfamily-SCOP Family A G protein-coupled receptor-like UniProtKB/Swiss-Prot, UniProtKB/TrEMBL
UniProt A6J2P8 ENTREZGENE, UniProtKB/TrEMBL
  GPR17_RAT UniProtKB/Swiss-Prot, ENTREZGENE


Nomenclature History
Date Current Symbol Current Name Previous Symbol Previous Name Description Reference Status
2006-11-20 Gpr17  G protein-coupled receptor 17      Symbol and Name status set to provisional 70820 PROVISIONAL