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The Vertebrate Trait Ontology (VT) is being developed as a collaborative effort between the Rat Genome Database at the Medical College of Wisconsin (RGD,, Mouse Genome Informatics at the Jackson Laboratory ( and the Animal QTL database at Iowa State University ( For more information about this vocabulary, or to request additions or changes, please contact us (

Term:surface structure morphology trait
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Accession:VT:0015066 term browser browse the term
Definition:Any measurable or observable characteristic related to the shape, structure, color or pattern of the organism subdivision which is the collection of anatomical structures on the body surface.

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coat/hair pigmentation trait term browser
Symbol Object Name Evidence Notes Source PubMed Reference(s) RGD Reference(s) Position
G Asip agouti signaling protein ISO Coat colour, agouti
Coat colour, saddle tan vs black-and-tan
OMIA PMID:8202545 PMID:12692166 PMID:15520882 PMID:15965787 PMID:17630272 More... NCBI chr 3:143,473,584...143,561,170
Ensembl chr 3:143,555,696...143,561,171
JBrowse link
G Defb14 defensin beta 14 ISO Coat colour, dominant black OMIA PMID:12692166 PMID:15649147 PMID:17483404 PMID:17947548 PMID:19197024 More... NCBI chr16:70,663,443...70,668,473
Ensembl chr16:70,663,333...70,666,540
JBrowse link
G Mc1r melanocortin 1 receptor ISO Coat colour, melanistic mask OMIA PMID:12692165 PMID:21353269 PMID:29932470 PMID:33672409 PMID:34751460 More... NCBI chr19:51,452,448...51,455,375
Ensembl chr19:51,453,239...51,454,192
JBrowse link
G Pmel premelanosome protein ISO Coat colour, merle OMIA PMID:330141 PMID:340189 PMID:418699 PMID:3099512 PMID:3998446 More... NCBI chr 7:1,138,202...1,148,735
Ensembl chr 7:1,138,586...1,148,746
JBrowse link
G Psmb7 proteasome 20S subunit beta 7 ISO Coat colour, harlequin OMIA PMID:3270445 PMID:3998446 PMID:18513894 PMID:21256207 PMID:31658272 More... NCBI chr 3:22,357,777...22,417,748
Ensembl chr 3:22,354,698...22,417,937
JBrowse link

Term paths to the root
Path 1
Term Annotations click to browse term
  vertebrate trait 2888
    organism subdivision trait 211
      surface structure trait 211
        surface structure morphology trait 211
          appendage morphology trait + 156
          beak morphology trait 0
          cloaca morphology trait 0
          ear morphology trait + 0
          eye morphology trait + 22
          integumentary system morphology trait + 38
          lip morphology trait + 0
          nose morphology trait + 0
          opercular flap morphology trait 0
          snout morphology trait + 0
paths to the root